Just what I have been trying, not saying its right or wrong just giving my experience.
For the Last couple months I have been trying my best to only feed every other day, I will admit that I am trying to get a couple random/picky fish onto other foods so I feed a little on the off days but definantly not allot of food and not targeting my rays. If I throw anything to the rays they are getting thrown some 10mm nls and they have actually started to love NLS allot when I feed it on the feeding day. At first they seemed a little cranky about not getting food daily but seem to have adjusted, if I feed some of the favorite food on the off day to the other fish the rays go a little crazy (but always do go crazy for talapia) while they can smell that food in the tank but settle down after. They are all fat and happy, My males have been trying to breed the females allot so I can't say anything about the eating of pregnant females yet, I keep watching them to see if they get less of a fat tail or back and have not seen it so far. I do feed a good bit on the days I do feed but not crazy amounts of food, and only feed once per day.
I know with actual fish I can judge overfeeding by looking at them and seeing how fat they are, just my though if the ray always has a huge back maybe they are being fed too much food as well and this later down the road can lead to a shorter life, I explained this in another thread that after I visited a local aquarium they were talking about deaths (they just lost their only Arapaima) and they found far more fat deposits and less life overall in which they attributed overfeeding as a cause to almost all fish that had died at the aquarium.