To be honest, all three fish need further studies to conclude if they are indeed separate species or just regional variations. While sp. coatzacoalcoas can display thinner red caudal band, other regional populations can display a thicker band. Paratheraps sp. coatzacoalcos are highly variable, which is interesting considering the short distribution range.
I have a hard time excepting current aquarium populations of guttulatus as they all said to originate from either Rio Sarabia and Rio Manzanita which are all part of the Coatzacoalcos basin. As you know P. guttulatus was originaly described from Lake Amatitlan Guatamala, quite a distance from Rio Coatzacoalcos.
The big question is- are they all the same fish? If they are then the name zonatus would no longer be valid as it would then become a junior synonym of guttulatus, many consider this a viable possibility.
With respect Günther's original guttulatus description is very loosely based and not much information since has come forward regarding Paratheraps forms from Pacific side Guatemala? Are guttulatus and zonatus the same? Perhaps, my gut feeling is that zonatus and sp. coatzacoalcos are the same fish at least?
Lee.
What you've said makes a lot of sense. The lack of clarity regarding the identity and validity of
guttulatus is a big problem, and more broadly, the entire group needs careful study to establish whether distinct species exist. For now I'm comfortable assuming that fish originating from the eastern drainage areas are in the
zonatus complex, which may include sp. coatzacoalcos.
Incidentally, the fish I posted above came from Cichlids of the Americas. I recently purchased some
P. zonatus 'Rio Jaltepec' from Jeff Rapps, and these individuals show more uniform red on the tail. Don't know the origin of the COA fish, but I'm guessing it comes from the Atlantic side as well.
Thanks for your response.
- Ian