Not much of anything that we do in our glass boxes imitate what takes place in the wild, so IMO whether a fish eats from the surface or below the surface doesn't have a whole lot of bearing in this discussion. (overfeeding) Some fish feel more comfortable eating from the surface, some seem to prefer sinking foods. Not that there's anything wrong with your method, but I wouldn't consider pellet types being part of the golden rule of feeding fish.
Not trying to nit pick, I completely agree with the rest of your points in that comment.
Feed less than you think necessary
A couple times a week skip feeding entirely
Feed smaller pellets
I see people feeding 10mm pellets to fish that are 6", I can only assume it's in the misguided belief that by doing so it will speed up the growth in their fish? Or feeding "high protein" foods, when their fish are fully mature. Again, it defies everything we know about the nutrient needs of adult cichlids, but it seems to be a common theme among many MONSTER fish keepers.
Sinking food also makes the fish work harder for their meal, it gets down in the gravel and fish need to do more "grazing" type feeding than just sucking down 10 floating pellets in 1 minute. Spreads the consumption out a bit. Same reason why I mention smaller pellets. The fish has to work harder and longer (expending energy) for an equivalent amount of energy (50 sinking 1mm pellets vs. 2 4.5mm floating pellets).
It has nothing to do with nutrition, and everything to do with spreading the feeding out.
It really depends on the fish, and how one feeds them. From a nutritional stand point I want my fish to consume pellet foods very quickly once those pellets enter the water column, to avoid water soluble nutrient loss. When using floating pellets, I feed only one or at the most two pellets at a time (depending on the size of the fish) moving from one tank to the next. I also break up their daily intake into two small feedings, one in the AM, and one in the PM. By doing so I am spreading out their overall consumption at any given time. Once a fish matures I only feed once a day at most (1 small feeding) with days within the week where no food is offered.
Also, as Jeff just mentioned using floating pellets I can control exactly what each fish consumes, with no concerns about any food getting lost within the substrate. A couple of my tanks contain rather large rocks/gravel, so in those tanks floaters make far more sense.
IMO the key point to be taken away from that previous paper that I linked to is this.
The nutrient content, energy content and digestibility of food resources all influence the overall quality of an animals diet.
Something that I have stated time & time again on this forum. Some of today's commercial foods are far more nutrient dense, have higher energy content, and far greater overall digestibility than foods manufactured 30 yrs ago. One can feed less, and overall nutrient wise get far more bang for your buck.
The trick is finding the sweet spot for each fish, based on their age, size, energy level, etc. Two fish of the same species, age & size, can require a vasty different amount of feed based soley on each fishes overall energy level in an aquarium. Some fish are laid back & mellow, some are hyper aggressive & never stop moving.
Learn something new every day. You're right Neil, I did not read the paper before posting and made the assumption that gut length was genotypic instead of phenotypic in nature. I apologize. No where though did I see the paper mention a phenotypic change of 50%. Can you point that out for me? I'll also point out that the study had a very small sample size (n = 4) for each area, which could lead to some different results if repeated.
Hey, no problem. I assumed that you hadn't read it when you posted. I don't believe that I stated that the paper linked to mentioned a phenotypic change of 50%. The 50% data came from an unpublished study performed years ago. I have no idea how many generations removed from the wild those "domestic" Tropheus were.
My point was simply that even among wild cichlids, they can quickly adapt to whatever food source is readily available.