Darker coloration, increased aggression associated with a specific object like a smooth rock, cleaning off rocks or smooth surfaces, collecting driftwood fragments and loose leaves. Their vents don't swell much until right before the spawn. They will flare and tail-whip at one another, but that is also common between same sex, so might not be a good indicator. Males sometimes nudge the side/vent area of females with their snout. In larger groups, increased aggression/territorial behavior between females is a good indicator that one or more is ready to breed. Their brood care is really interesting and entertaining, good luck!
Hope that helps,
Ed