But most of their lives, those rheophiles spend time in quite fast, highly oxygenated conditions....The average Orinoco River flow rate in S America is 1.37 million gallons per hour, at 0 ppm nitrate so maybe its true...... capturing any Geo from the river to keep in a 4 ft tank with a canister filter as pumpage is a tad ridiculous.
Those numbers sound impressive, and are impressive, but...let's do some math.
A quick search of the stats for the Orinoco gives a vast range of average widths and depths, largely depending upon the season (wet vs dry). The bottom extreme of this range...the narrowest average width...was shown as about a kilometer (1000m or about 3300 feet). Likewise, the shallowest average depth was claimed as 50 meters (about 165 feet). That gives us an average cross-sectional area of well over a half million square feet. Since the bottom of the river slopes downward from the shore to the deepest part and then back up again to the opposite shore, let's divide that area in half...one quarter million square feet.
As that 1.37 million gallons of water per hour hurtles downstream, it is spread over a quarter million square feet of cross-sectional area. A quick calculation seems to show that the flow rate per square foot is something around 5 gallons per hour per square foot.
Of course this is too simplistic; there will be huge variations from area to area. So multiply our number by ten, or even by 100...it's still somewhat underwhelming. Set up a tank, of any length, with a depth and a height of one foot. Slap on a canister or a HOB that moves 500 gallons per hour (not unrealistic). Extend the intake to one end of the tank and the return to the other end. You have a created a microcosm that approaches, matches or even exceeds the monumental flow rate of the mighty Orinoco river.
Please, prove me wrong. I was never fully behind the idea that flow rates in nature were so far in excess of those in our tanks...but now I'm shocked to see how feeble they are in practical terms. I don't care about the huge numbers needed to describe the overall amount of water rushing to the sea; I'm thinking more about the actual amount of water flowing around the fish where they are situated.
On top of that, I can't help but compare this to my observations of trout in their natural environment. Everybody "knows" that trout "need" fast-flowing, highly-aerated water...but the fact that they often live in small, shallow, clear-water streams makes them a prime candidate for "fish-watching". No need to flop around in the water with them; it's fairly simple to find an elevated bank or other dry, comfortable perch from which to observe them. Surprise! They don't spend most, or even much, of their time fighting a raging current. They almost always lurk in a quiet section of slack water, in the lee of a boulder or other obstruction, from which they launch their feeding attacks on passing prey items.
Usually, the biggest/oldest/most dominant fish are found in places like beaver ponds or still-water pools along the stream. The big guys are no dummies; they, like most animals, follow the path of least resistance to conserve as much energy as possible.
Certainly, they need high oxygen levels, as the water in which they live has been thoroughly oxygenated in riffles, rapids and shallows...but they find much less energy-intensive places to rest and enjoy all that wonderful oxygen.