I've seen poor growth or stunting result from sickness or stress at the fry or juvie stage. Protein or feed levels that are way off from their requirements can also be a factor (including way too high according to some studies I've seen). Sometimes you just get a weak individual that seems to struggle and not grow well from the start (might be simple genetics, but also biochemistry issues, like hormones that don't fire as they're supposed to, whether due to environmental, nutritional, or other factors).
Also, some ingredients/food sources that are digestible at later stages are not so much at early or initial fry stages when their digestive process is still in development, and some foods that are good at early stages are less beneficial as the fish develops. This aspect gets complicated, so rather than try to research and figure out all the science I just go with personal experience of what works for me.
I've also had runts that eventually overcame whatever their early weakness was and later turn into nice big adults.
Also, some ingredients/food sources that are digestible at later stages are not so much at early or initial fry stages when their digestive process is still in development, and some foods that are good at early stages are less beneficial as the fish develops. This aspect gets complicated, so rather than try to research and figure out all the science I just go with personal experience of what works for me.
I've also had runts that eventually overcame whatever their early weakness was and later turn into nice big adults.