if you use a UGF in conjunction with a HOB (for example, an ac110) the bacteria will establish itself in the HOB rather than the UGF as it provides a better habitat for the bacteria (higher flow rate. more oxygen etc) at which point the UGF isnt really doing anything initially. and then once the media in the HOB is established, the bacteria will continue to colonize elsewhere as the ammonia levels in the tank rise.
as the system runs, the UGF will trap rotting debris under the tank raising the amount of available ammonia for the bacteria to convert, and the system will continue to balance itself out to the amount of "food" for the bacteria. but now your biological filtration is running based on the amount of crap under the plates, instead of the daily bio-load of the fish themselves.
the system will run just as well with only the HOB, and the biological filtration will balance out with the bio-load of the fish, rather than the added ammonia from the detrius trapped under the UGF.
(feel free to correct me if im mistaken)