I have to weigh in on the other side of the fence on this one. I do not think the pigmentation that some rays exhibit on the bellies have anything to do with the yolk sac. Not all rays have them, but all rays have yolk sacs. The location of the belly spot also is not always in the same location and the yolk sac remnant on every pup I have ever seen is in the same place. You might expect some type of scar- which we do not see- but not a dark pigmented spot. Also, you do not see any type of yolk sac scarring on any other rays or sharks to the best of my knowledge, not to mention reptiles that also are nourished through a yolk sac.
I think it is just pigment. Rays have spots on the top of their discs, why can't they on the underside.
I think it is just pigment. Rays have spots on the top of their discs, why can't they on the underside.