I agree with nc_nutcase. I've kept JDs for over 35 years, and when mine get stressed they go light not dark. If you look closely at the pic I posted, my dark JD is in the process of spawning with the Gold JD just below it. In my experience stressed fish don't tend to spawn readily. While I agree that dark substrate can cause a fish to darken, mine are on a white sand substrate, so the dark background theory doesn't seem to matter with some JDs. It seems that some individuals are just naturally darker than the norm. Hypermelanism is not unusual in many animals and fish.
Mark