looks like you've ruled out the stress from other fish then. it seems that its eaither something your not testing for or something thats returned back to normal when you do test.
is it possible to turn the drip off and leave it to your filters if the drip is where you think the problem is ?
your talking about getting rid of alot of stock to try fix the issues but if you have already have a successfull tank for reference and have moved females to another tank that runs off the same larger system why dont you just make that third tank/system independant using a duplicate of the successfull tanks filtration ? hopefully no further problems and then you can just move the adults back to the main tank after the pups are born leaving the pups undisturbed and you can keep the main tank as it is.
is it possible to turn the drip off and leave it to your filters if the drip is where you think the problem is ?
your talking about getting rid of alot of stock to try fix the issues but if you have already have a successfull tank for reference and have moved females to another tank that runs off the same larger system why dont you just make that third tank/system independant using a duplicate of the successfull tanks filtration ? hopefully no further problems and then you can just move the adults back to the main tank after the pups are born leaving the pups undisturbed and you can keep the main tank as it is.
I don't know how much of a difference it takes before its an issue. I'm sure much less for newborns then established rays.